Biodiversity
The aquatic flora is limited more by the habitat requirements of the individual species than by any geographical restriction of range. Most species are found almost throughout Britain wherever suitable habitats occur, and the restriction of some species to certain parts of the country reflects the localisation of suitable habitats.
In terms of their distribution in the British Isles the aquatic flora can be divided into five main groups; widespread & common; widespread & local; northern species; southern species; and rare species.
The Review gives special prominence to the vertebrates particularly in relation to their associations with the main nutrient types of standing water.
Open waters are more likely to be altered by man's activity than completely destroyed, and few open water sites in Britain have disappeared entirely. The modification of open waters can be so profound however that they become valueless to nature conservation, and in extreme cases (e.g. heavily polluted rivers) they may be virtually lifeless. Pollution, including eutrophication, is the major modifying factor affecting open waters, and chemical and biotic tests for different forms of pollution play a part in the comparative evaluation of sites. Since most sources of pollution originate at points within the catchment remote from the waterbody, naturalness of the catchment, as well as intactness of the water-body itself, is important. In the case of oligo-trophic sites the effects of slight pollution or enrichment are more drastic and the degree of intactness demanded of the catchment is much greater than that demanded of eutrophic sites. Fortunately most oligotrophic waters are in the relatively unpopulated uplands, but aerial applications of fertilisers may pose a threat to them.
The hydrology of open waters may also be affected by man's activities. Many natural lakes have been converted to storage reservoirs in which rapid and large- scale fluctuations in water level cause severe impoverishment of the littoral and sublittoral fauna and flora. The greater and more rapid the drawdown in a reservoir the less is it likely to be of value for conservation, and in this respect hydro-electric reservoirs are usually of less value than water supply reservoirs. In rivers, the hydrology is modified by the construction of locks, weirs and dams, and by regulation of discharge, embankment and straightening as well as water abstraction and effluent discharge. Most of these modifications lead to a deterioration in the structural and biological diversity of river systems.
In lowland Britain increased recreational activity on many open waters is destroying much aquatic wildlife and the presence of the more destructive forms of recreation on a water-body has counted against that site in the overall assessment.
Natural water-bodies are normally preferable to those artificially constructed, particularly in relation to the stratigraphical history depicted in their sediments, and artificial water-bodies have only been chosen where no natural alternative exists. However, some types of water-body such as the shallow fenland meres (e.g. the former Whittlesea Mere) are no longer found in a natural state in Britain and, since many of the interesting invertebrate species are now extinct in this country, this ecosystem is irretrievably lost and artificial waters now form the nearest alternative. In many parts of lowland Britain, particularly in the chalk and limestone areas, there are very few natural standing water-bodies, and artificial types, such as Tring Reservoirs and the Cotswold Water Park gravel pits, are habitats not previously found in Britain, yet have high conservation value. Many rivers, particularly the chalk streams of southern England, which are often thought of as a natural part of the British scene, are highly modified and only maintained in their present form by intensive management involving dredging, weed-cutting, removal of coarse fish, and stocking with trout. They nevertheless represent a diverse habitat not found in a natural state in Britain.
In spite of the modifications to many British freshwater stystems, some of these, including a high proportion of Scottish Highland lochs and streams, are probably the habitats least changed by man in inland Britain.
Structural diversity
This is an extremely important criterion in the assessment of open waters, since, probably more than for any other formation, chemistry and morphology are the most critical factors controlling the flora and fauna. Diversity of shoreline is particularly important in the case of lakes and, since the coarser sediments only occur in exposed situations, larger lakes are more diverse than smaller ones. Similarly, the presence of deep water gives additional diversity.