The hominoids include both the ape line—the
Pongidae—and the line leading to man—the Hominidae. The
Pongidae are represented today by the great apes— the
chimpanzee, the gorilla and the orang-utang. The Hominidae include
the fossil Australopithecus, fossil man, now extinct, and living
man—Homo sapiens. Although this advance first took
place in the New World, there was no further progress in this
hemisphere beyond the level of the New World monkeys. But in the
Old World this new type developed not only into the Old World
monkeys, but gave rise to a progressive branch which advanced from
the pithecoid or monkey stage to the hominoid stage. This is
'man-like' and includes the gibbons, the great apes and man
today.
It is now believed by a number of investigators
that this original, unspecialized, monkey-like type gave rise not
to one hominoid line which divided into men and apes at a very late
stage, but into two main streams: the Pongidae, leading to the
great apes, and the Hominidae leading to man. As long ago as the
Oligocene there is evidence of these two divergent types of
hominoid.
It was for a long time believed that the
divergence took place only 14 million years ago or thereabouts, and
that a Miocene African ape, Proconsul, which was relatively
unspecialized and had not developed. the anatomical modifications
for swinging in trees known as brachiation, was the common ancestor
of men and apes.
More recent theories classify Proconsul as
belonging to a most interesting and widely dispersed genus of
tree-apes known as Dryopithecus, which was the culmination of the
pongid line and which separated from the hominids 20 million years
before Proconsul. It is thought that Proconsul or a similar type
may have given rise to the chimpanzee and gorilla, and that another
type of Dryopithecus may have been the ancestor of the orang-utang.
The other forms of this genus all became extinct.
The hominid line never modified in a brachiating
direction and never developed the powerful canine teeth found in
all pongids including Proconsul. We know that at the same time as
Proconsul there existed at least two distinctive hominids which are
almost certainly either the immediate ancestors of Australopithecus
(the precursor of man) or was actually the first known
Australopithecus. These hominids are known as Ramapithecus and
Kenyapithecus, the latter discovered by Leaky in 1962. Ramapithecus
has been found in India, Africa and Europe.
It has been suggested that there may be a
connection between this hominid line and a very unspecialized
little monkey-like type, Propliopithecus, which lived in Egypt 34
million years ago and had the pattern of teeth and jaw
characteristic of the later hominids. At roughly the same time
there was, also in Egypt, another type with well marked canines,
Aegyptopithecus, which may indicate the ancestral type of the
Pongidae.